When playing levels, you can choose to use keyboard controls, or a gamepad.

WASD: move

Space: jump

Shift: run/grab

Esc/P: pause

Mouse scroll or Comma/Period: cycle through scrollable items in the UI

In the editor, a mouse and keyboard are used together for building levels.

Left click: use current tool

Mouse scroll: change zoom

WASD: pan

Tab: toggle play/edit

Esc/Q: select tool, cancel

F: player start position hotkey

X: hide/show editor toolbars

E: eraser hotkey

R: rotate current sprite/tile (if it can be rotated)

Z: undo

Y: redo

Delete: delete selection

Control: hold and drag selection to copy